Table 4.

Gene editing studies in hemophilia

Gene targetedCell typeGene editingNuclease deliveryDonorResults in recipient model
Hemophilia B      
F949
Li, 2011 
Hepatocytes Gene correction ZFN via plasmid and AAV delivery system DNA Correction of FIX in neonatal HB mouse model 
F950
Anguela, 2013 
Hepatocytes Gene correction ZFN via plasmid and AAV delivery system DNA Correction of FIX in adult HB mouse model 
F951
Lyu, 2018 
iPSCs Gene addition Cas9 via plasmid electroporation Plasmid FIX secreted in transplanted HB mouse 
F952
Ramaswamy, 2018 
iPSCs Gene addition Cas 9 plasmid nucleofection DNA Correction of FIX in HLCs in HB mouse 
F9353
Morishige, 2020 
iPSCs Gene correction Cas9 plasmid nucleofection DNA Detection of transfected HLCs in HB mouse 
F954
Stephens, 2019 
Stem cells Gene addition Cas9 plasmid and Ad platform Plasmid Correction of FIX in HB mice 
F955
Wang, 2020 
Hepatocytes Gene addition Cas9 plasmid and AAV delivery system Plasmid Correction of FIX in adult and neonatal HB mice 
F956
Ma, 2022 
ESCs Gene addition Cas9 plasmid electroporation Plasmid Detection of FIX at the RNA and protein levels 
F957
Lee, 2022 
Hepatocyte Gene addition Cas9 plasmid and AAV delivery system Plasmid Detection of FIX in HB mouse model 
F958
He, 2022 
Hepatocyte Gene addition Cas9 plasmid and AAV delivery system Plasmid Correction of FIX in adult and neonatal HB mice 
Hemophilia A      
F959
Sharma, 2015 
Hepatocytes Gene correction ZFN via plasmid and AAV delivery system DNA Correction of FVIII in HA mouse model 
F860
Park, 2015 
iPSCs Gene correction Cas9 via plasmid electroporation None FVIII activity restored in HA mouse model 
F861
Hu, 2019 
iPSCs Gene correction Cas9 via plasmid nucleofection ssODNs FVIII activity restored in HA mouse model 
F862
Park, 2019 
iPSCs Gene addition Cas9 via RNP electroporation DNA FVIII secreted in the iPSCs of patients with HA 
F863
Sung, 2019 
iPSCs Gene addition Cas9 via plasmid electroporation Plasmid FVIII secreted in the iPSCs of patients with HA 
F864
Chen, 2019 
Hepatocytes Gene addition Cas9 via plasmid and AAV delivery system Plasmid FVIII secreted in HA mouse hepatocytes 
F865
Zhang, 2019 
Hepatocytes Gene addition Cas9 via plasmid and AAV delivery system Plasmid FVIII secreted in HA mouse hepatocytes 
F866
Ramamurthy, 2021 
Placental cells Gene addition Cas9 via plasmid delivery system Plasmid FVIII secreted in human placental cells 
F867
Cho, 2021 
iPSCs Gene correction Cas9 via plasmid delivery system Plasmid FVIII secreted in the iPSCs of patients with HA 
F868
Hu, 2023 
iPSCs Gene correction Cas9 via plasmid delivery system Plasmid FVIII secreted in the iPSCs of patients with HA 
Gene targetedCell typeGene editingNuclease deliveryDonorResults in recipient model
Hemophilia B      
F949
Li, 2011 
Hepatocytes Gene correction ZFN via plasmid and AAV delivery system DNA Correction of FIX in neonatal HB mouse model 
F950
Anguela, 2013 
Hepatocytes Gene correction ZFN via plasmid and AAV delivery system DNA Correction of FIX in adult HB mouse model 
F951
Lyu, 2018 
iPSCs Gene addition Cas9 via plasmid electroporation Plasmid FIX secreted in transplanted HB mouse 
F952
Ramaswamy, 2018 
iPSCs Gene addition Cas 9 plasmid nucleofection DNA Correction of FIX in HLCs in HB mouse 
F9353
Morishige, 2020 
iPSCs Gene correction Cas9 plasmid nucleofection DNA Detection of transfected HLCs in HB mouse 
F954
Stephens, 2019 
Stem cells Gene addition Cas9 plasmid and Ad platform Plasmid Correction of FIX in HB mice 
F955
Wang, 2020 
Hepatocytes Gene addition Cas9 plasmid and AAV delivery system Plasmid Correction of FIX in adult and neonatal HB mice 
F956
Ma, 2022 
ESCs Gene addition Cas9 plasmid electroporation Plasmid Detection of FIX at the RNA and protein levels 
F957
Lee, 2022 
Hepatocyte Gene addition Cas9 plasmid and AAV delivery system Plasmid Detection of FIX in HB mouse model 
F958
He, 2022 
Hepatocyte Gene addition Cas9 plasmid and AAV delivery system Plasmid Correction of FIX in adult and neonatal HB mice 
Hemophilia A      
F959
Sharma, 2015 
Hepatocytes Gene correction ZFN via plasmid and AAV delivery system DNA Correction of FVIII in HA mouse model 
F860
Park, 2015 
iPSCs Gene correction Cas9 via plasmid electroporation None FVIII activity restored in HA mouse model 
F861
Hu, 2019 
iPSCs Gene correction Cas9 via plasmid nucleofection ssODNs FVIII activity restored in HA mouse model 
F862
Park, 2019 
iPSCs Gene addition Cas9 via RNP electroporation DNA FVIII secreted in the iPSCs of patients with HA 
F863
Sung, 2019 
iPSCs Gene addition Cas9 via plasmid electroporation Plasmid FVIII secreted in the iPSCs of patients with HA 
F864
Chen, 2019 
Hepatocytes Gene addition Cas9 via plasmid and AAV delivery system Plasmid FVIII secreted in HA mouse hepatocytes 
F865
Zhang, 2019 
Hepatocytes Gene addition Cas9 via plasmid and AAV delivery system Plasmid FVIII secreted in HA mouse hepatocytes 
F866
Ramamurthy, 2021 
Placental cells Gene addition Cas9 via plasmid delivery system Plasmid FVIII secreted in human placental cells 
F867
Cho, 2021 
iPSCs Gene correction Cas9 via plasmid delivery system Plasmid FVIII secreted in the iPSCs of patients with HA 
F868
Hu, 2023 
iPSCs Gene correction Cas9 via plasmid delivery system Plasmid FVIII secreted in the iPSCs of patients with HA 

Ad, adenoviral vector; ESCs, embryonic stem cells; RNP, ribonucleoprotein; ssODNs, single-stranded oligodeoxynucleotides.

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